Seems like you need these relationships:
MouseColony|Male::UID = MatingRecord::M UID
MouseColony|Female::UID = MatingRecord::F UID
MouseColony|Male and MouseColony|Female would be two Tutorial: What are Table Occurrences? with MouseColony as the same data source table for both.
For an explanation of the notation that I am using, see the first post of: Common Forum Relationship and Field Notations Explained
This worked perfectly! Thank you for the rapid response!